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Defender Squads are built from Defenders. There are four Defender Squads, one for each Map Location.
You choose defenders to include in a given location's squad, and then when you're defending your base in that location, you can summon those defenders with Defender Posts. There are a number of different slots that can be unlocked for each base, each matching a specific kind of combat, like melee, shotgun, sniper, and so on.
Each defender has one randomly chosen bonus for each five levels he gains. These bonuses only apply only to the defender himself, not the team.